Genome Biology | Full text | Fish eye = development

From: Subject: Genome Biology | Full text | Fish eye development Date: Wed, 5 May 2004 17:43:23 -0700 MIME-Version: 1.0 Content-Type: multipart/related; boundary="----=_NextPart_000_0021_01C432C8.769DD640"; type="text/html" X-MimeOLE: Produced By Microsoft MimeOLE V6.00.2800.1106 This is a multi-part message in MIME format. ------=_NextPart_000_0021_01C432C8.769DD640 Content-Type: text/html; charset="utf-8" Content-Transfer-Encoding: quoted-printable Content-Location: http://genomebiology.com/2000/1/4/reports/0070 =EF=BB=BF Genome Biology | Full text | Fish eye = development

5-May-2004

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=20 Issue=20 4

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Liu=20 = K

Paper report
Fish=20 eye development

The importance of the lens in eye development has = been=20 demonstrated using reciprocal transplantations between an eyed=20 surface-dwelling fish and an eyeless cave fish of the same = species.

Central role for the lens = in cave fish=20 eye degeneration.
Y = Yamamoto and WR=20 Jeffrey Science = 2000,=20 289:631-633 [Read the = abstract on=20 PubMed]

Subject areas: = Development,=20 Evolution, Model organisms

Reported by: Karen Liu

The electronic version of this article is the = complete one and=20 can be found online at: http://genomebiol= ogy.com/2000/1/4/reports/0070

Report = date		13 September 2000
Published		25 October 2000

Genome Biology 2000,=20 1:reports0070

=C2=A9 2000=20 BioMed Central Ltd

Outline

Significance and context

Abstract<= /A>

Significa= nce=20 and context

Key=20 results

Links=

Reporter'= s=20 comments

Table=20 of links

In Mexico, there are over 30 varieties of the = fish=20 Astyanax mexicanus. Some populations of these fish live in = surface=20 waters while others live in unlit caves. The cave-dwelling = varieties are=20 eyeless, providing a unique opportunity to study the evolution of = a=20 developmental process in a single species, rather than in = distantly=20 related species. Using an eyed surface-dweller and an eyeless=20 cave-dweller, Yamamoto and Jeffrey are able to show that an = inductive=20 signal from the lens influences eye formation.

In the surface-dwelling fish, eyes form by a = process quite=20 similar to that in other vertebrates. First, the optic vesicle = buds out=20 from the brain and contacts the overlying ectoderm, which forms = the lens=20 precursor. The optic vesicle folds to become the optic cup, while = the lens=20 develops and becomes internalized. The optic cup then = differentiates into=20 the retina and the ectoderm overlying the lens becomes the cornea. = In the=20 cave fish, the lens and optic cup develop normally for the first = 24 hours;=20 after this, however, the lens cells undergo programmed cell death = or=20 apoptosis. The cornea and iris do not form and the retina is = extremely=20 disorganized. Growth of the eye stops and photoreceptor cells = never=20 develop. The rudimentary eye eventually sinks back and is covered = by=20 skin.

The authors proposed two hypotheses for eye = degeneration in=20 the cave fish. In one, lens apoptosis is responsible for eye = degeneration.=20 In the other, the optic cup, which is diminished in the cave fish, = produces insufficient amounts of some necessary signal, leading to = the=20 deterioration of the eye and lens.

Outline

Key results

Abstract<= /A>

Significa= nce=20 and context

Key=20 results

Links=

Reporter'= s=20 comments

Table=20 of links

Yamamoto and Jeffrey transplanted embryonic = lenses from=20 eyed fish into the optic cups of eyeless fish and vice versa. They = found=20 that a lens from a surface fish could induce normal-looking eyes = in the=20 blind cave fish, whereas the reciprocal transplantation caused a=20 degeneration in the eye of the surface fish. In each case, the=20 unmanipulated eye on the other side served as a control. To = determine=20 whether programmed cell death was responsible for this = deterioration, the=20 authors looked for apoptosis in the donor lenses. They found that = the cave=20 fish lens still underwent apoptosis when implanted in a surface = fish=20 embryo and the resulting eye was small and disorganized. In the = opposite=20 experiment, the surface fish lens did not undergo apoptosis and = was able=20 to stimulate normal eye development in the blind cave fish host, = including=20 differentiation of a lens, retina, iris and cornea. The restored = cave fish=20 eye also expressed the transcription factor Pax6, which is = ordinarily=20 present in the lens and corneal epithelia but is not detected in=20 unmanipulated cave fish eye precursors. Yamamoto and Jeffrey then = labeled=20 the eyed fish donor tissue with a fluorescent dye prior to = implantation to=20 determine whether the rescued eye parts were from host or donor = tissues.=20 When they sectioned the embryos, only the lens was fluorescently = labeled,=20 revealing that the rescued eye parts were from the cave fish = tissues. This=20 showed that the implanted lens from the surface fish was capable = of=20 inducing the development of eye parts that were normally lost in = the cave=20 fish.

The authors also used molecular markers of = retinal=20 development to investigate the organization of the restored eye. = Pax6,=20 Prox1, proliferating cell nuclear antigen (PCNA) and rhodopsin = mark=20 different cell types within the retina. Pax6, Prox1 and PCNA were = all=20 expressed in the cave fish eye; however, the retinal layers were=20 disorganized and very few cells expressed rhodopsin, which marks = the rod=20 cells. In the restored eye, the retina developed into distinct, = organized=20 layers, indicating that normal patterning, growth and = differentiation of=20 the eye can be induced in the cave fish.

Links

A comment on this paper appeared in the same = issue of=20 Science and is available to subscribers (Pennisi E, Science289:552b). A = Supplementary figure to = Science289:631-633=20 can be accessed freely from the Science = website.

Reporter's comments

This study shows quite clearly that the lens can = play an=20 important role in patterning the eye. When the surface fish lens = is=20 implanted into the blind cave fish, normal eye development is = restored,=20 though it is unclear whether the restored eye is actually = functional. The=20 authors note that there are at least 30 Astyanax cave fish=20 populations in north-east Mexico. Their results apply only to the = Pach=C3=B3n=20 cave fish used in their study. The other varieties of blind cave = fish may=20 have evolved by different mechanisms and further comparisons of = these fish=20 may provide insight into the evolution of developmental=20 processes.

Table of links

Science

Pennisi E, Science289:552b

Supplementary figure to=20 Science289:631-633

Assumptions that = are made=20 about each paper that is the subject of a report, unless = otherwise=20 specified:

The full text and figures are available = only to=20 subscribers of the journal, but are available over the = internet from=20 the journal's website. The paper itself is abstracted by = PubMed.=20 There is no supplementary = material.

Published by

=C2=A9 1999-2004 BioMed Central Ltd = unless otherwise=20 stated < info@genomebiology.com=20 > Terms and=20 conditions

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client info flag = (1=3Don|0=3Doff) --*/ var __utmdn=3D"auto"; /*-- (auto|none|domain) set the domain = name for cookies --*/ var __utmhash=3D"on"; /*-- (on|off) unique domain hash for = cookies --*/ var __utmgifpath=3D"/__utm.gif"; /*-- set the web path to the __utm.gif = file --*/ var __utmtimeout=3D"1800"; /*-- set the inactive session timeout = in seconds --*/ /*-------------------------------------------------- UTM Campaign Tracking Settings --------------------------------------------------*/ var __utmctm=3D1; /*-- set campaign tracking module = (1=3Don|0=3Doff) --*/ var __utmcto=3D"15768000"; /*-- set the campaign timeout in = seconds (6 month default) --*/ var __utmccn=3D"utm_campaign"; /*-- campaign name --*/ var __utmcmd=3D"utm_medium"; /*-- campaign medium = (cpc|cpm|link|email|organic) --*/ var __utmcsr=3D"utm_source"; /*-- campaign source --*/ var __utmctr=3D"utm_term"; /*-- campaign term/keyword --*/ var __utmcct=3D"utm_content"; /*-- campaign content --*/ var __utmcid=3D"utm_id"; /*-- campaign id number --*/ var __utmcno=3D"utm_nooverride"; /*-- don't override campaign = information--*/ /*--- Auto/Organic Sources and Keywords ---*/ var __utmOsr =3D new Array(); var __utmOkw =3D new Array(); __utmOsr[0] =3D "google"; __utmOkw[0] =3D "q"; __utmOsr[1] =3D "yahoo"; __utmOkw[1] =3D "p"; __utmOsr[2] =3D "msn"; __utmOkw[2] =3D "q"; __utmOsr[3] =3D "aol"; __utmOkw[3] =3D "query"; __utmOsr[4] =3D "lycos"; __utmOkw[4] =3D "query"; __utmOsr[5] =3D "ask"; __utmOkw[5] =3D "q"; __utmOsr[6] =3D "altavista"; __utmOkw[6] =3D "q"; __utmOsr[7] =3D "search"; __utmOkw[7] =3D "q"; __utmOsr[8] =3D "netscape"; __utmOkw[8] =3D "query"; __utmOsr[9] =3D "earthlink"; __utmOkw[9] =3D "q"; __utmOsr[10] =3D "cnn"; __utmOkw[10] =3D "query"; __utmOsr[11] =3D "looksmart"; __utmOkw[11] =3D "key"; __utmOsr[12] =3D "about"; __utmOkw[12] =3D "terms"; __utmOsr[13] =3D "excite"; __utmOkw[13] =3D "qkw"; __utmOsr[14] =3D "mamma"; __utmOkw[14] =3D "query"; __utmOsr[15] =3D "alltheweb"; __utmOkw[15] =3D "q"; __utmOsr[16] =3D "gigablast"; __utmOkw[16] =3D "q"; __utmOsr[17] =3D "voila"; __utmOkw[17] =3D "kw"; __utmOsr[18] =3D "virgilio"; __utmOkw[18] =3D "qs"; __utmOsr[19] =3D "teoma"; __utmOkw[19] =3D "q"; /*--- Auto/Organic Keywords to Ignore ---*/ var __utmOno =3D new Array(); //__utmOno[0] =3D "urchin"; //__utmOno[1] =3D "urchin.com"; //__utmOno[2] =3D "www.urchin.com"; /*--- Referral domains to Ignore ---*/ var __utmRno =3D new Array(); //__utmRno[0] =3D ".urchin.com"; /*-------------------------------------------------- Don't modify below this point --------------------------------------------------*/ var __utmf,__utmdh,__utmd,__utmdom=3D"",__utmu,__utmjv=3D"-",__utmfns, = __utmns=3D0,__utmr=3D"-"; var __utmcfno=3D0; if (!__utmf) { var __utma,__utmb,__utmc; var __utmexp=3D"",__utms=3D"",__utmst=3D0,__utmlf=3D0; /*-- get useful information --*/ __utmdh =3D __utmSetDomain(); __utma =3D document.cookie.indexOf("__utma=3D"+__utmdh); __utmb =3D document.cookie.indexOf("__utmb=3D"+__utmdh); __utmc =3D document.cookie.indexOf("__utmc=3D"+__utmdh); __utmu =3D Math.round(Math.random() * 2147483647); __utmd =3D new Date(); __utmst =3D Math.round(__utmd.getTime()/1000); if (__utmdn && __utmdn !=3D "") { __utmdom =3D " = domain=3D"+__utmdn+";"; } if (__utmtimeout && __utmtimeout !=3D "") { __utmexp =3D new Date(__utmd.getTime()+(__utmtimeout*1000)); __utmexp =3D " expires=3D"+__utmexp.toGMTString()+";"; } /*-- grab cookies from the commandline --*/ __utms =3D document.location.search; if (__utms && __utms !=3D "" && __utms.indexOf("__utma=3D") >=3D 0) { __utma =3D __utmGetCookie(__utms,"__utma=3D","&"); __utmb =3D __utmGetCookie(__utms,"__utmb=3D","&"); __utmc =3D __utmGetCookie(__utms,"__utmc=3D","&"); if (__utma !=3D "-" && __utmb !=3D "-" && __utmc !=3D "-") __utmlf = =3D 1; else if (__utma !=3D "-") __utmlf =3D = 2; } /*-- based on the logic set cookies --*/ if (__utmlf =3D=3D 1) {=20 document.cookie=3D"__utma=3D"+__utma+"; path=3D/; expires=3DSun, = 18 Jan 2038 00:00:00 GMT;"; document.cookie=3D"__utmb=3D"+__utmb+"; path=3D/;"+__utmexp; document.cookie=3D"__utmc=3D"+__utmc+"; path=3D/;"; } else if (__utmlf =3D=3D 2) {=20 __utma =3D __utmFixA(__utms,"&",__utmst);=20 document.cookie=3D"__utma=3D"+__utma+"; path=3D/; expires=3DSun, = 18 Jan 2038 00:00:00 GMT;"; document.cookie=3D"__utmb=3D"+__utmdh+"; path=3D/;"+__utmexp; document.cookie=3D"__utmc=3D"+__utmdh+"; path=3D/;" __utmfns=3D1; } else if (__utma >=3D 0 && __utmb >=3D 0 && __utmc >=3D 0) {=20 document.cookie=3D"__utmb=3D"+__utmdh+"; = path=3D/;"+__utmexp+__utmdom; } else if (__utma >=3D0) {=20 __utma =3D __utmFixA(document.cookie,";",__utmst);=20 document.cookie=3D"__utma=3D"+__utma+"; path=3D/; expires=3DSun, = 18 Jan 2038 00:00:00 GMT;"+__utmdom; document.cookie=3D"__utmb=3D"+__utmdh+"; = path=3D/;"+__utmexp+__utmdom; document.cookie=3D"__utmc=3D"+__utmdh+"; path=3D/;"+__utmdom; __utmfns=3D1; } else if (__utma < 0 && __utmb < 0 && __utmc < 0) {=20 __utma =3D __utmCheckUTMI(__utmd);=20 if (__utma =3D=3D "-") __utma =3D = __utmdh+"."+__utmu+"."+__utmst+"."+__utmst+"."+__utmst+".1";=20 else __utma =3D __utmdh+"."+__utma; document.cookie=3D"__utma=3D"+__utma+"; path=3D/; expires=3DSun, = 18 Jan 2038 00:00:00 GMT;"+__utmdom; document.cookie=3D"__utmb=3D"+__utmdh+"; = path=3D/;"+__utmexp+__utmdom; document.cookie=3D"__utmc=3D"+__utmdh+"; path=3D/;"+__utmdom; __utmfns=3D1; } else { __utma =3D = __utmdh+"."+__utmu+"."+__utmst+"."+__utmst+"."+__utmst+".1"; document.cookie=3D"__utma=3D"+__utma+"; path=3D/; expires=3DSun, = 18 Jan 2038 00:00:00 GMT;"+__utmdom; document.cookie=3D"__utmb=3D"+__utmdh+"; = path=3D/;"+__utmexp+__utmdom; document.cookie=3D"__utmc=3D"+__utmdh+"; path=3D/;"+__utmdom; __utmfns=3D1; } __utmSetInfo(); __utmf =3D 1; } function __utmSetInfo() { var __utmp; var __utmi =3D new Image(1,1); var __utmsrc =3D __utmgifpath+"?"; var loc =3D document.location; __utmr =3D document.referrer; if (!__utmr || __utmr =3D=3D "") { __utmr =3D "-"; }=20 else {=20 __utmp =3D __utmr.indexOf(document.domain);=20 if ((__utmp >=3D 0) && (__utmp <=3D 8)) { __utmr =3D "0"; } if (__utmr.indexOf("[") =3D=3D 0 && __utmr.lastIndexOf("]") =3D=3D = (__utmr.length-1)) { __utmr =3D "-"; } } __utmsrc +=3D "utmn=3D"+__utmu; if (__utmfsc && __utmfns) {__utmsrc +=3D __utmGetClientInfo(); } if (__utmctm) {__utmsrc +=3D __utmSetCampaignInfo(); } __utmsrc +=3D "&utmr=3D"+__utmr+"&utmp=3D"+loc.pathname+loc.search; __utmi.src =3D __utmsrc; __utmi.onload =3D function() { __utmVoid(); } return 0; } function __utmVoid() { return; } function __utmSetCampaignInfo() { var __utmcc =3D ""; var __utmtmp =3D "-"; var __utmtmp2 =3D "-"; var __utmnoover =3D 0; var __utmcsc =3D 0; var __utmcnc =3D 0; var __utmi =3D 0; if (!__utmcto || __utmcto =3D=3D "") { __utmcto =3D "15768000"; } var __utmcx =3D new Date(__utmd.getTime()+(__utmcto*1000)); __utmcx =3D " expires=3D"+__utmcx.toGMTString()+";"; var __utmx =3D document.location.search; var __utmz =3D __utmGetCookie(__utmx,"__utmz=3D","&"); if (__utmz !=3D "-") { document.cookie=3D"__utmz=3D"+__utmz+"; = path=3D/;"+__utmcx+__utmdom; return ""; } __utmz =3D document.cookie.indexOf("__utmz=3D"+__utmdh); if (__utmz > -1) { __utmz =3D __utmGetCookie(document.cookie,"__utmz=3D",";"); } else { __utmz =3D "-"; } /*--- check for campaign id or campaign source ---*/ __utmtmp =3D __utmGetCookie(__utmx,__utmcid+"=3D","&"); __utmtmp2 =3D __utmGetCookie(__utmx,__utmcsr+"=3D","&"); if ((__utmtmp !=3D "-" && __utmtmp !=3D "") || (__utmtmp2 !=3D "-" = && __utmtmp2 !=3D "")) {=20 if (__utmtmp !=3D "-" && __utmtmp !=3D "") { __utmcc +=3D "utmcid=3D"+__utmtmp; if (__utmtmp2 !=3D "-" && __utmtmp2 !=3D "") __utmcc +=3D = "|utmcsr=3D"+__utmtmp2; } else { if (__utmtmp2 !=3D "-" && __utmtmp2 !=3D "") __utmcc +=3D = "utmcsr=3D"+__utmtmp2; } __utmtmp =3D __utmGetCookie(__utmx,__utmccn+"=3D","&");=20 if (__utmtmp !=3D "-" && __utmtmp !=3D "") __utmcc +=3D = "|utmccn=3D"+__utmtmp;=20 else __utmcc +=3D = "|utmccn=3D(not set)"; __utmtmp =3D __utmGetCookie(__utmx,__utmcmd+"=3D","&");=20 if (__utmtmp !=3D "-" && __utmtmp !=3D "") __utmcc +=3D = "|utmcmd=3D"+__utmtmp; else __utmcc +=3D = "|utmcmd=3D(not set)"; __utmtmp =3D __utmGetCookie(__utmx,__utmctr+"=3D","&");=20 if (__utmtmp !=3D "-" && __utmtmp !=3D "") {=20 __utmcc +=3D "|utmctr=3D"+__utmtmp; } else {=20 __utmtmp =3D __utmGetOrganic(1); if (__utmtmp !=3D "-" && __utmtmp !=3D "") __utmcc +=3D = "|utmctr=3D"+__utmtmp;=20 } __utmtmp =3D __utmGetCookie(__utmx,__utmcct+"=3D","&");=20 if (__utmtmp !=3D "-" && __utmtmp !=3D "") __utmcc +=3D = "|utmcct=3D"+__utmtmp; __utmtmp =3D __utmGetCookie(__utmx,__utmcno+"=3D","&");=20 if (__utmtmp =3D=3D "1") __utmnoover =3D 1; /*--- if previous campaign is set and no override is set return = ---*/ if (__utmz !=3D "-" && __utmnoover =3D=3D 1) return ""; } /*--- check for organic ---*/ if (__utmcc =3D=3D "-" || __utmcc =3D=3D "") { __utmcc =3D __utmGetOrganic();=20 /*--- if previous campaign is set and organic no override term is = found return ---*/ if (__utmz !=3D "-" && __utmcfno =3D=3D 1) return ""; } /*--- check for referral ---*/ if (__utmcc =3D=3D "-" || __utmcc =3D=3D "") { if (__utmfns =3D=3D 1) __utmcc =3D __utmGetReferral();=20 /*--- if previous campaign is set and referral no override term = is found return ---*/ if (__utmz !=3D "-" && __utmcfno =3D=3D 1) return ""; } /*--- set default if z is not yet set ---*/ if (__utmcc =3D=3D "-" || __utmcc =3D=3D "") { if (__utmz =3D=3D "-" && __utmfns =3D=3D 1) { __utmcc =3D = "utmccn=3D(direct)|utmcsr=3D(direct)|utmcmd=3D(none)"; } if (__utmcc =3D=3D "-" || __utmcc =3D=3D "") return ""; } /*--- check if campaign is already set and if it's the same ---*/ if (__utmz !=3D "-") {=20 __utmi =3D __utmz.indexOf("."); if (__utmi > -1) __utmi =3D __utmz.indexOf(".",__utmi+1); if (__utmi > -1) __utmi =3D __utmz.indexOf(".",__utmi+1); if (__utmi > -1) __utmi =3D __utmz.indexOf(".",__utmi+1); __utmtmp =3D __utmz.substring(__utmi + 1,__utmz.length); if (__utmtmp.toLowerCase() =3D=3D __utmcc.toLowerCase()) __utmcsc = =3D 1;=20 __utmtmp =3D __utmz.substring(0,__utmi); if ((__utmi =3D __utmtmp.lastIndexOf(".")) > -1) { __utmtmp =3D __utmtmp.substring(__utmi+1,__utmtmp.length); __utmcnc =3D (__utmtmp*1); } } /*--- set the cookie ---*/ if (__utmcsc =3D=3D 0 || __utmfns =3D=3D 1) { __utmtmp =3D __utmGetCookie(document.cookie,"__utma=3D",";"); if ((__utmi=3D__utmtmp.lastIndexOf(".")) > 9) { __utmns =3D __utmtmp.substring(__utmi+1,__utmtmp.length); __utmns =3D (__utmns*1); } __utmcnc++; if (__utmns =3D=3D 0) __utmns =3D 1; = document.cookie=3D"__utmz=3D"+__utmdh+"."+__utmst+"."+__utmns+"."+__utmcn= c+"."+__utmcc+"; path=3D/; "+__utmcx+__utmdom; } /*--- set the new campaign flag ---*/ if (__utmcsc =3D=3D 0 || __utmfns =3D=3D 1) return "&utmcn=3D1"; else return "&utmcr=3D1"; } function __utmGetReferral() { if (__utmr =3D=3D "0" || __utmr =3D=3D "" || __utmr =3D=3D "-") = return "";=20 var __utmi=3D0; var __utmhn; var __utmkt; /*-- get the hostname of the referral --*/ if ( (__utmi =3D __utmr.indexOf("://")) < 0) return ""; __utmhn =3D __utmr.substring(__utmi+3,__utmr.length); if (__utmhn.indexOf("/") > -1) { __utmkt =3D = __utmhn.substring(__utmhn.indexOf("/"),__utmhn.length); if (__utmkt.indexOf("?") > -1) { __utmkt =3D __utmkt.substring(0,__utmkt.indexOf("?")); } __utmhn =3D __utmhn.substring(0,__utmhn.indexOf("/")); } __utmhn =3D __utmhn.toLowerCase(); for (var ii=3D0;ii<__utmRno.length;ii++) { if (( __utmi=3D__utmhn.indexOf(__utmRno[ii].toLowerCase())) > -1 = && __utmhn.length =3D=3D (__utmi+__utmRno[ii].length)) { __utmcfno =3D = 1; break; } } if (__utmhn.indexOf("www.") =3D=3D 0) { __utmhn =3D __utmhn.substring(4,__utmhn.length); } return = "utmccn=3D(referral)|utmcsr=3D"+__utmhn+"|"+"utmcct=3D"+__utmkt+"|utmcmd=3D= referral"; } function __utmGetOrganic(termonly) { if (__utmr =3D=3D "0" || __utmr =3D=3D "" || __utmr =3D=3D "-") = return "";=20 var __utmi=3D0; var __utmhn; var __utmkt; /*-- get the hostname of the referral --*/ if ( (__utmi =3D __utmr.indexOf("://")) < 0) return ""; __utmhn =3D __utmr.substring(__utmi+3,__utmr.length); if (__utmhn.indexOf("/") > -1) { __utmhn =3D __utmhn.substring(0,__utmhn.indexOf("/")); } for (var ii=3D0;ii<__utmOsr.length;ii++) { if (__utmhn.indexOf(__utmOsr[ii]) > -1) { if ( (__utmi =3D __utmr.indexOf("?"+__utmOkw[ii]+"=3D")) > -1 = ||=20 (__utmi =3D __utmr.indexOf("&"+__utmOkw[ii]+"=3D")) > -1) = { __utmkt =3D = __utmr.substring(__utmi+__utmOkw[ii].length+2,__utmr.length); if ( (__utmi =3D __utmkt.indexOf("&")) > -1) { __utmkt =3D __utmkt.substring(0,__utmi); } for (var yy=3D0;yy<__utmOno.length;yy++) { if (__utmOno[yy].toLowerCase() =3D=3D = __utmkt.toLowerCase()) { __utmcfno =3D 1; break; } } if (termonly) { return __utmkt; } else { return = "utmccn=3D(organic)|utmcsr=3D"+__utmOsr[ii]+"|"+"utmctr=3D"+__utmkt+"|utm= cmd=3Dorganic"; } } } } return ""; } function __utmGetClientInfo() { var = __utmtmp=3D"-",__utmsr=3D"-",__utmsa=3D"-",__utmsc=3D"-",__utmbs=3D"-",__= utmul=3D"-"; var __utmje=3D1,__utmce=3D1,__utmtz=3D0; if (self.screen) {=20 __utmsr =3D screen.width+"x"+screen.height; __utmsa =3D screen.availWidth+"x"+screen.availHeight; __utmsc =3D screen.colorDepth+"-bit"; } else if (self.java) { var __utmjk =3D java.awt.Toolkit.getDefaultToolkit(); var __utmjksize =3D __utmjk.getScreenSize(); =20 __utmsr =3D __utmjksize.width+"x"+__utmjksize.height; }=20 if( typeof( window.innerWidth ) =3D=3D 'number' ) { __utmbs =3D window.innerWidth+"x"+window.innerHeight; } else {=20 if (document.documentElement &&=20 (document.documentElement.offsetHeight || = document.documentElement.offsetWidth ) ) { __utmbs =3D = document.documentElement.offsetWidth+"x"+document.documentElement.offsetH= eight; } else if (document.body && (document.body.offsetWidth || = document.body.offsetHeight) ) { __utmbs =3D = document.body.offsetWidth+"x"+document.body.offsetHeight; }=20 } for (var i=3D5;i>=3D0;i--) { var __utmtmp =3D "";=20 document.write(__utmtmp); if (__utmjv !=3D "-") break; } if (navigator.language) { __utmul =3D = navigator.language.toLowerCase(); } else if (navigator.browserLanguage) { __utmul =3D = navigator.browserLanguage.toLowerCase(); } __utmje =3D navigator.javaEnabled()?1:0; if (document.cookie.indexOf("__utmb=3D") < 0) { __utmce =3D "0"; } if (document.cookie.indexOf("__utmc=3D") < 0) { __utmce =3D "0"; } __utmtz =3D __utmd.getTimezoneOffset(); __utmtz =3D __utmTZConvert(__utmtz); __utmtmp =3D""; __utmtmp +=3D = "&utmsr=3D"+__utmsr+"&utmsa=3D"+__utmsa+"&utmsc=3D"+__utmsc+"&utmbs=3D"+_= _utmbs; __utmtmp +=3D = "&utmul=3D"+__utmul+"&utmje=3D"+__utmje+"&utmce=3D"+__utmce+"&utmtz=3D"+_= _utmtz+"&utmjv=3D"+__utmjv; return __utmtmp; } function __utmLinker(__utmlink) { var = __utmlp,__utmi,__utmi2,__utmta=3D"-",__utmtb=3D"-",__utmtc=3D"-",__utmtz=3D= "-"; if (__utmlink && __utmlink !=3D "") {=20 if (document.cookie) { __utmta =3D = __utmGetCookie(document.cookie,"__utma=3D"+__utmdh,";"); __utmtb =3D = __utmGetCookie(document.cookie,"__utmb=3D"+__utmdh,";"); __utmtc =3D = __utmGetCookie(document.cookie,"__utmc=3D"+__utmdh,";"); __utmtz =3D = __utmGetCookie(document.cookie,"__utmz=3D"+__utmdh,";"); __utmlp =3D = "__utma=3D"+__utmta+"&__utmb=3D"+__utmtb+"&__utmc=3D"+__utmtc+"&__utmz=3D= "+__utmtz; } if (__utmlp) { if (__utmlink.indexOf("?") <=3D -1) { document.location =3D = __utmlink+"?"+__utmlp; } else { document.location =3D __utmlink+"&"+__utmlp; } } else { document.location =3D __utmlink; } } } function __utmGetCookie(__utmclist,__utmcname,__utmcsep) { if (!__utmclist || __utmclist =3D=3D "") return "-"; if (!__utmcname || __utmcname =3D=3D "") return "-"; if (!__utmcsep || __utmcsep =3D=3D "") return "-"; var __utmi, __utmi2, __utmi3, __utmtc=3D"-"; __utmi =3D __utmclist.indexOf(__utmcname); __utmi3 =3D __utmcname.indexOf("=3D")+1; if (__utmi > -1) {=20 __utmi2 =3D __utmclist.indexOf(__utmcsep,__utmi); if (__utmi2 < 0) = { __utmi2 =3D __utmclist.length; } __utmtc =3D __utmclist.substring((__utmi+__utmi3),__utmi2);=20 } return __utmtc; } function __utmSetDomain() { if (!__utmdn || __utmdn =3D=3D "" || __utmdn =3D=3D "none") { __utmdn = =3D ""; return 1; } if (__utmdn =3D=3D "auto") { var __utmdomain =3D document.domain; if (__utmdomain.substring(0,4) =3D=3D "www.") { __utmdomain =3D __utmdomain.substring(4,__utmdomain.length); } __utmdn =3D __utmdomain; } if (__utmhash =3D=3D "off") return 1; return __utmHash(__utmdn); } function __utmHash(__utmd) { if (!__utmd || __utmd =3D=3D "") return 1; var __utmhash=3D0, __utmg=3D0; for (var i=3D__utmd.length-1;i>=3D0;i--) { var __utmc =3D parseInt(__utmd.charCodeAt(i));=20 __utmhash =3D ((__utmhash << 6) & 0xfffffff) + __utmc + (__utmc << = 14); if ((__utmg =3D __utmhash & 0xfe00000) !=3D 0) __utmhash =3D = (__utmhash ^ (__utmg >> 21)); } return __utmhash; } function __utmFixA(__utmcs,__utmsp, __utmst) { if (!__utmcs || __utmcs =3D=3D "") return "-"; if (!__utmsp || __utmsp =3D=3D "") return "-"; if (!__utmst || __utmst =3D=3D "") return "-"; var __utmt =3D __utmGetCookie(__utmcs,"__utma=3D",__utmsp); var __utmlt=3D0; var __utmi=3D0; if ((__utmi=3D__utmt.lastIndexOf(".")) > 9) { __utmns =3D __utmt.substring(__utmi+1,__utmt.length); __utmns =3D (__utmns*1)+1; __utmt =3D __utmt.substring(0,(__utmi)); if ((__utmi =3D __utmt.lastIndexOf(".")) > 7) { __utmlt =3D __utmt.substring(__utmi+1,__utmt.length); __utmt =3D __utmt.substring(0,(__utmi)); } if ((__utmi =3D __utmt.lastIndexOf(".")) > 5) { __utmt =3D __utmt.substring(0,(__utmi)); } __utmt +=3D "."+__utmlt+"."+__utmst+"."+__utmns; } return __utmt; } function __utmCheckUTMI(__utmd) { var __utm1A =3D new Array(); var = __utmlst=3D0,__utmpst=3D0,__utmlvt=3D0,__utmlu=3D0,__utmi=3D0,__utmpi=3D0= ; var __utmap =3D "-"; var __utmld =3D ""; var __utmt2; var __utmt =3D document.cookie; while((__utmi =3D __utmt.indexOf("__utm1=3D")) >=3D 0) { __utm1A[__utm1A.length] =3D = __utmGetCookie(__utmt,"__utm1=3D",";"); __utmt =3D __utmt.substring(__utmi+7,__utmt.length); } if (__utm1A.length) { var __utmcts =3D Math.round(__utmd.getTime()/1000); var __utmlex =3D " expires=3D"+__utmd.toGMTString()+";"; __utmt =3D document.cookie;=20 if ((__utmi =3D __utmt.lastIndexOf("__utm3=3D")) >=3D 0) { __utmlst =3D __utmt.substring(__utmi,__utmt.length); __utmlst =3D __utmGetCookie(__utmlst,"__utm3=3D",";"); } if ((__utmi =3D __utmt.lastIndexOf("__utm2=3D")) >=3D 0) { __utmpst =3D __utmt.substring(__utmi,__utmt.length); __utmpst =3D __utmGetCookie(__utmpst,"__utm2=3D",";"); } for (var i=3D0;i<__utm1A.length;i++) { __utmt =3D __utm1A[i]; if ((__utmi =3D __utmt.lastIndexOf(".")) >=3D 0) { __utmt2 =3D (__utmt.substring(0,__utmi))*1; __utmt =3D (__utmt.substring(__utmi+1,__utmt.length))*1; if (__utmlvt =3D=3D 0 || __utmt < __utmlvt) {=20 __utmlvt =3D __utmt; __utmlu =3D __utmt2; } } } if (__utmlvt && __utmlst) {=20 if (!__utmpst || __utmpst > __utmlst) __utmpst =3D __utmlst; __utmap =3D = __utmlu+"."+__utmlvt+"."+__utmpst+"."+__utmlst+".2";=20 } else if (__utmlvt) {=20 if (!__utmpst || __utmpst > __utmcts) __utmpst =3D __utmcts; __utmap =3D = __utmlu+"."+__utmlvt+"."+__utmpst+"."+__utmcts+".2"; } __utmld =3D __utmt =3D document.domain; __utmi=3D__utmpi=3D0; while((__utmi =3D __utmt.indexOf(".",__utmpi+1)) >=3D 0) { if (__utmpi>0) __utmld =3D = __utmt.substring(__utmpi+1,__utmt.length); __utmld =3D " domain=3D"+__utmld+";";=20 document.cookie=3D"__utm1=3D1; path=3D/;"+__utmlex+__utmld; document.cookie=3D"__utm2=3D1; path=3D/;"+__utmlex+__utmld; document.cookie=3D"__utm3=3D1; path=3D/;"+__utmlex+__utmld; __utmpi=3D__utmi; } document.cookie=3D"__utm1=3D1; path=3D/;"+__utmlex; document.cookie=3D"__utm2=3D1; path=3D/;"+__utmlex; document.cookie=3D"__utm3=3D1; path=3D/;"+__utmlex; } return __utmap; } function __utmTZConvert(__utmmz) { var __utmhr=3D0,__utmmn=3D0,__utmsg=3D'+'; if (__utmmz && __utmmz !=3D "") { if (__utmmz <=3D 0) {__utmsg=3D'+'; __utmmz*=3D-1; } else {__utmsg=3D'-'; __utmmz*=3D1; } __utmhr =3D Math.floor((__utmmz/60));=20 __utmmn =3D Math.floor((__utmmz%60));=20 } if (__utmhr < 10) __utmhr =3D "0"+__utmhr; if (__utmmn < 10) __utmmn =3D "0"+__utmmn; return __utmsg+__utmhr+__utmmn; } ------=_NextPart_000_0021_01C432C8.769DD640--