Genome Biology | Full text | Fish eye = development

From: Subject: Genome Biology | Full text | Fish eye development Date: Wed, 5 May 2004 17:43:23 -0700 MIME-Version: 1.0 Content-Type: multipart/related; boundary="----=_NextPart_000_0021_01C432C8.769DD640"; type="text/html" X-MimeOLE: Produced By Microsoft MimeOLE V6.00.2800.1106 This is a multi-part message in MIME format. ------=_NextPart_000_0021_01C432C8.769DD640 Content-Type: text/html; charset="utf-8" Content-Transfer-Encoding: quoted-printable Content-Location: http://genomebiology.com/2000/1/4/reports/0070 =EF=BB=BF Genome Biology | Full text | Fish eye = development

5-May-2004

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=20 Issue=20 4

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Liu=20 = K

Paper report
Fish=20 eye development

The importance of the lens in eye development has = been=20 demonstrated using reciprocal transplantations between an eyed=20 surface-dwelling fish and an eyeless cave fish of the same = species.

Central role for the lens = in cave fish=20 eye degeneration.
Y = Yamamoto and WR=20 Jeffrey Science = 2000,=20 289:631-633 [Read the = abstract on=20 PubMed]

Subject areas: = Development,=20 Evolution, Model organisms

Reported by: Karen Liu

The electronic version of this article is the = complete one and=20 can be found online at: http://genomebiol= ogy.com/2000/1/4/reports/0070

Report = date		13 September 2000
Published		25 October 2000

Genome Biology 2000,=20 1:reports0070

=C2=A9 2000=20 BioMed Central Ltd

Outline

Significance and context

Abstract<= /A>

Significa= nce=20 and context

Key=20 results

Links=

Reporter'= s=20 comments

Table=20 of links

In Mexico, there are over 30 varieties of the = fish=20 Astyanax mexicanus. Some populations of these fish live in = surface=20 waters while others live in unlit caves. The cave-dwelling = varieties are=20 eyeless, providing a unique opportunity to study the evolution of = a=20 developmental process in a single species, rather than in = distantly=20 related species. Using an eyed surface-dweller and an eyeless=20 cave-dweller, Yamamoto and Jeffrey are able to show that an = inductive=20 signal from the lens influences eye formation.

In the surface-dwelling fish, eyes form by a = process quite=20 similar to that in other vertebrates. First, the optic vesicle = buds out=20 from the brain and contacts the overlying ectoderm, which forms = the lens=20 precursor. The optic vesicle folds to become the optic cup, while = the lens=20 develops and becomes internalized. The optic cup then = differentiates into=20 the retina and the ectoderm overlying the lens becomes the cornea. = In the=20 cave fish, the lens and optic cup develop normally for the first = 24 hours;=20 after this, however, the lens cells undergo programmed cell death = or=20 apoptosis. The cornea and iris do not form and the retina is = extremely=20 disorganized. Growth of the eye stops and photoreceptor cells = never=20 develop. The rudimentary eye eventually sinks back and is covered = by=20 skin.

The authors proposed two hypotheses for eye = degeneration in=20 the cave fish. In one, lens apoptosis is responsible for eye = degeneration.=20 In the other, the optic cup, which is diminished in the cave fish, = produces insufficient amounts of some necessary signal, leading to = the=20 deterioration of the eye and lens.

Outline

Key results

Abstract<= /A>

Significa= nce=20 and context

Key=20 results

Links=

Reporter'= s=20 comments

Table=20 of links

Yamamoto and Jeffrey transplanted embryonic = lenses from=20 eyed fish into the optic cups of eyeless fish and vice versa. They = found=20 that a lens from a surface fish could induce normal-looking eyes = in the=20 blind cave fish, whereas the reciprocal transplantation caused a=20 degeneration in the eye of the surface fish. In each case, the=20 unmanipulated eye on the other side served as a control. To = determine=20 whether programmed cell death was responsible for this = deterioration, the=20 authors looked for apoptosis in the donor lenses. They found that = the cave=20 fish lens still underwent apoptosis when implanted in a surface = fish=20 embryo and the resulting eye was small and disorganized. In the = opposite=20 experiment, the surface fish lens did not undergo apoptosis and = was able=20 to stimulate normal eye development in the blind cave fish host, = including=20 differentiation of a lens, retina, iris and cornea. The restored = cave fish=20 eye also expressed the transcription factor Pax6, which is = ordinarily=20 present in the lens and corneal epithelia but is not detected in=20 unmanipulated cave fish eye precursors. Yamamoto and Jeffrey then = labeled=20 the eyed fish donor tissue with a fluorescent dye prior to = implantation to=20 determine whether the rescued eye parts were from host or donor = tissues.=20 When they sectioned the embryos, only the lens was fluorescently = labeled,=20 revealing that the rescued eye parts were from the cave fish = tissues. This=20 showed that the implanted lens from the surface fish was capable = of=20 inducing the development of eye parts that were normally lost in = the cave=20 fish.

The authors also used molecular markers of = retinal=20 development to investigate the organization of the restored eye. = Pax6,=20 Prox1, proliferating cell nuclear antigen (PCNA) and rhodopsin = mark=20 different cell types within the retina. Pax6, Prox1 and PCNA were = all=20 expressed in the cave fish eye; however, the retinal layers were=20 disorganized and very few cells expressed rhodopsin, which marks = the rod=20 cells. In the restored eye, the retina developed into distinct, = organized=20 layers, indicating that normal patterning, growth and = differentiation of=20 the eye can be induced in the cave fish.

Links

A comment on this paper appeared in the same = issue of=20 Science and is available to subscribers (Pennisi E, Science289:552b). A = Supplementary figure to = Science289:631-633=20 can be accessed freely from the Science = website.

Reporter's comments

This study shows quite clearly that the lens can = play an=20 important role in patterning the eye. When the surface fish lens = is=20 implanted into the blind cave fish, normal eye development is = restored,=20 though it is unclear whether the restored eye is actually = functional. The=20 authors note that there are at least 30 Astyanax cave fish=20 populations in north-east Mexico. Their results apply only to the = Pach=C3=B3n=20 cave fish used in their study. The other varieties of blind cave = fish may=20 have evolved by different mechanisms and further comparisons of = these fish=20 may provide insight into the evolution of developmental=20 processes.

Table of links

Science

Pennisi E, Science289:552b

Supplementary figure to=20 Science289:631-633

Assumptions that = are made=20 about each paper that is the subject of a report, unless = otherwise=20 specified:

The full text and figures are available = only to=20 subscribers of the journal, but are available over the = internet from=20 the journal's website. The paper itself is abstracted by = PubMed.=20 There is no supplementary = material.

Published by

=C2=A9 1999-2004 BioMed Central Ltd = unless otherwise=20 stated < info@genomebiology.com=20 > Terms and=20 conditions

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